R¹, R² and R³ are each a hydrogen atom;

Q¹ represents a saturated or unsaturated, 5- or 6-membered cyclic hydrocarbon group which may be substituted, a saturated or unsaturated, 5- to 7-membered heterocyclic group which may be substituted, a saturated or unsaturated, bicyclic or tricyclic condensed hydrocarbon group which may be substituted, or a saturated or unsaturated, bicyclic or tricyclic condensed heterocyclic group which may be substituted;

Q² represents a single bond, a linear or branched alkylene group having 1 to 6 carbon atoms, a linear or branched alkenylene group having 2 to 6 carbon atoms, a linear or branched alkynylene group having 2 to 6 carbon atoms, a saturated or unsaturated, 5- or 6-membered divalent cyclic hydrocarbon group which may be substituted, a saturated or unsaturated, 5- to 7-membered divalent heterocyclic group which may be substituted, a saturated or unsaturated, divalent bicyclic or tricyclic condensed hydrocarbon group which may be substituted, or a saturated or unsaturated, divalent bicyclic or tricyclic condensed heterocyclic group which may be substituted;

R⁴ represents a substituted or unsubstituted 5-membered heterocyclic group;

Q⁴ represents an aryl group which may be substituted, an arylalkenyl group which may be substituted, an arylalkynyl group which may be substituted, a heteroaryl group which may be substituted, a heteroarylalkenyl group which may be substituted, a saturated or unsaturated, bicyclic or tricyclic condensed hydrocarbon group which may be substituted, or a saturated or unsaturated, bicyclic or tricyclic condensed heterocyclic group which may be substituted;

T⁰ represents a carbonyl or thiocarbonyl group; and

T¹ represents a carbonyl group, sulfonyl group, group —C(═O)—C(═O)—N(R′)—, group —C(═S)—C(═O)—N(R′)—, group —C(═O)—C(═S)—N(R′)—, group —C(═S)—C(═S)—N(R′)— (wherein R′ represents a hydrogen atom, hydroxyl group, alkyl group or alkoxy group), group —C(═O)-A¹-N(R″)— (wherein A¹ represents an alkylene group which has 1 to 5 carbon atoms and may be substituted, and R″ represents a hydrogen atom, hydroxyl group, alkyl group or alkoxy group, group), —C(═O)—NH—, group —C(═S)—NH—, group —C(═O)—NH—NH—, group —C(═O)-A²-C(═O)— (wherein A² represents a single bond or alkylene group having 1 to 5 carbon atoms), group —C(═O)-A³-C(═O)—NH— (wherein A³ represents an alkylene group having 1 to 5 carbon atoms), group —C(═O)—C(═NOR^a)—N(R^b), group —C(═S)—C(═NOR^a)—N(R^b)— (wherein R^a represents a hydrogen atom, alkyl group or alkanoyl group, and R^b represents a hydrogen atom, hydroxyl group, alkyl group or alkoxy group), group —C(═O)—N═N—, group —C(═S)—N═N—, group —C(═NOR^c)—C(═O)—N(R^d)— (wherein R^c represents a hydrogen atom, alkyl group, alkanoyl group, aryl group or aralkyl group, and R^d represents a hydrogen atom, hydroxyl group, alkyl group or alkoxy group), group —C(═N—N(R^e)(R^f))—C(═O)—N(R^g)— (wherein R^e and R^f each independently represent a hydrogen atom, alkyl group, alkanoyl group or alkyl(thiocarbonyl) group, and R^g represents a hydrogen atom, hydroxyl group, alkyl group or alkoxy group), group —C(═O)—NH—C(═O)—, group —C(═S)—NH—C(═O)—, group —C(═O)—NH—C(═S)—, group —C(═S)—NHC(═S)—, group —C(═O)—NH—SO₂—, group —SO₂—NH—, group —C(═NCN)—NH—C(═O)—, group —C(═S)—C(═O)— or thiocarbonyl group; a salt thereof.